Concept:
During prophase I of meiosis, homologous chromosomes pair with each other in a process called synapsis. Prophase I is divided into five stages:
• Leptotene
• Zygotene (Zygonema)
• Pachytene
• Diplotene
• Diakinesis
The synaptonemal complex is a protein structure that forms between homologous chromosomes and facilitates pairing and recombination.
Step 1: Understanding when synaptonemal complex forms.
During:
\[
\text{Zygotene (Zygonema)}
\]
homologous chromosomes begin pairing through synapsis.
At this stage:
\[
\text{Synaptonemal complex begins to form}
\]
Therefore:
\[
\boxed{\text{Synaptonemal complex begins to form during Zygonema}}
\]
Step 2: Understanding the role of Pachytene.
In Pachytene:
• Synapsis becomes complete
• Crossing over occurs
Thus, the complex is already fully formed by Pachytene.
Step 3: Eliminating the incorrect options.
• Option (A): Anaphase involves chromosome separation.
• Option (B): Metaphase involves chromosome alignment at the equatorial plate.
• Option (C): Pachytene is the stage where crossing over occurs after synaptonemal complex formation.
Hence, the correct answer is Zygonema.
\[
\boxed{\text{Zygonema} \Rightarrow \text{Beginning of synaptonemal complex formation}}
\]