Step 1: Carl Correns worked on the four o'clock plant, Mirabilis jalapa, which shows variegated leaves with green, white and pale branches.
Step 2: He found that the colour of the offspring depended only on the branch that provided the female parent (the seed-bearing branch), not on the pollen source, showing maternal, non-Mendelian inheritance.
Step 3: This pattern is explained by plastids, which carry their own DNA and are transmitted mainly through the cytoplasm of the egg cell, not through pollen.
Step 4: Mitochondria, nucleus and Golgi bodies were not the structures involved in this classic leaf-variegation study, so plastids is correct.