Step 1: Understanding the Concept:
Gravitropism is the coordinated process of plant growth in response to gravity.
Roots exhibit positive gravitropism (growing downward), which is mediated by specialized cells that can sense gravitational pull.
Step 2: Detailed Explanation:
- Gravity sensing occurs in specialized cells called statocytes located in the center of the root cap (the columella).
- Statocytes contain heavy, starch-filled organelles called amyloplasts (specifically referred to as statoliths in the context of gravitropism).
- Because of their high density relative to the surrounding cytoplasm, amyloplasts sediment to the lowest part of the cell in response to gravity.
- This sedimentation exerts physical pressure on the endoplasmic reticulum and cytoskeleton, triggering a signaling cascade that redistributes the plant hormone auxin.
- The resulting asymmetric distribution of auxin inhibits cell elongation on the lower side of the root, causing it to curve downward.
- Other plastids like chloroplasts (photosynthetic), chromoplasts (pigmented), and gerontoplasts (decaying plastids) do not contain dense starch granules and do not function in gravity sensing.
Step 3: Final Answer:
The gravity-sensing plastids that direct downward root growth are amyloplasts.