Step 1: Understanding anther structure.
The anther wall consists of four layers: epidermis, endothecium, middle layers, and tapetum.
Step 2: Role of endothecium.
The endothecium develops fibrous thickenings that generate tension during drying, which leads to anther dehiscence.
Step 3: Eliminating other layers.
Epidermis provides protection.
Tapetum nourishes developing pollen grains.
Middle layers degenerate during development.
Step 4: Conclusion.
Therefore, the endothecium is responsible for anther dehiscence.