Statement I is true.
\textit{Daphnia}, a planktonic crustacean, exhibits cyclomorphosis — a seasonal variation in body shape.
Statement II is incorrect.
In a thermally stratified lake of temperate regions, during winter, the upper layer is actually hypolimnion, not epilimnion.
Statement III is true.
A niche represents the functional role an organism plays in its environment, including interactions, habitat use, and resource consumption.
Statement IV is incorrect.
Bergmann’s rule relates body size to environmental temperature, not metabolic rate. It states that endothermic animals tend to be larger in colder climates.
Hence, the correct statements are I and III.