Step 1: Understanding the Concept:
Crassulacean Acid Metabolism (CAM) is a specialized photosynthetic adaptation found in succulent plants growing in arid environments.
It involves a temporal separation of carbon dioxide capture and assimilation to minimize water loss.
Step 2: Detailed Explanation:
Let us evaluate both statements to understand the physiological traits of CAM plants:
Assertion A Analysis: CAM plants possess thick, fleshy, water-storing tissues (succulence) which allow them to store substantial water reserves to survive extreme drought.
However, this adaptation does not enhance their photosynthetic efficiency.
In fact, CAM plants have extremely low photosynthetic efficiency and slow growth rates compared to C3 and C4 plants.
Succulence is an adaptation to maximize water-use efficiency and survival, not photosynthetic rate or efficiency.
Therefore, Assertion A is false.
Reason R Analysis: In CAM plants, stomata open at night to capture \(\text{CO}_2\), which is fixed by PEP carboxylase into malate.
This malate must be stored inside the large central vacuole of the mesophyll cells as malic acid to prevent cellular acidification.
During the day, stomata close, and the stored malic acid is transported out of the vacuole and decarboxylated to release \(\text{CO}_2\) for Rubisco.
Because the plant can only capture as much \(\text{CO}_2\) at night as it can store as malate in its vacuoles, the overall photosynthetic capacity is strictly limited by the vacuolar storage volume.
Therefore, Reason R is true.
Step 3: Final Answer:
Assertion A is false but Reason R is true, which corresponds to option (D).