Concept:
Gynogenesis is the production of haploids from the female gametophyte (megaspore/egg cell). The success of this process depends on the developmental stage of the female tissues.
Step 1: Evaluating the Assertion (The Error).
Assertion (A) is incorrect. Gynogenic haploids are developed from female gametophytic tissues. The "optimum stage of male gametophytic development" would be relevant for androgenesis (anther/pollen culture), not gynogenesis.
Step 2: Evaluating the Reason.
Reason (R) is correct. For the successful induction of haploid embryos from female tissues, the megaspore or embryo sac needs to be at a specific maturity level, generally ranging from the late uninucleate stage up to the mature trinucleate/organized embryo sac stage.
Step 3: Conclusion.
Because the Assertion misidentifies the gametophyte source for gynogenesis while the Reason correctly identifies the developmental stages of the female megaspore, option (4) is the correct choice.