Step 1: Understanding the Concept:
Plasmodesmata are microscopic channels that traverse the cell walls of plant cells, enabling transport and communication between them.
Step 2: Detailed Explanation:
A plasmodesma contains a central tubule called a desmotubule.
During cell division, as the new cell plate forms, tubules of the Endoplasmic Reticulum (ER) become trapped within the developing wall.
These trapped ER tubules are compressed into a narrow structure that forms the desmotubule.
The desmotubule is continuous with the cortical endoplasmic reticulum of both adjacent cells, creating a shared endomembrane network (the symplast).
Step 3: Final Answer:
The desmotubule in plasmodesmata is formed from the Endoplasmic Reticulum network.