(A) NADP reductase: This enzyme is located on the stroma side of the thylakoid membrane. However, it is associated with Photosystem I (PS I). According to standard texts (e.g., NCERT), stroma lamellae membranes lack PS II and NADP reductase. This is why only cyclic photophosphorylation occurs there. Thus, placing it in stroma lamellae is a mismatch.
(B) PEPcase: Found in the cytosol of Mesophyll cells in C$_4$ plants. Correct.
(C) RuBisCO: Found in the Bundle sheath cells of C$_4$ plants (and mesophyll of C$_3$). Correct.
(D) ATP synthetase: Found in the thylakoid membrane of the Chloroplast. Correct.
Step 2: Conclusion:
Option (A) is the mismatched pair because the enzyme is absent in stroma lamellae.