Step 1: Match each enzyme with its application.
Phytase → improves mineral availability (4).
Xylanase → used in paper and pulp processing (1).
Laccase → involved in delignification (2).
Bromelain → helps reduce gluten complexes (3).
Step 2: Compare with options.
This matches option (B): P-4, Q-1, R-2, S-3.
Step 3: Conclusion.
Thus, option (B) gives the correct matching.